This is the A-to-Z story of Andrew and Zelda - a pair that almost wasn’t - and all that happened from the day they met. Andrew has always been a secret romantic … not above crooning to Celine Dion while driving to work, with dreams of finding “the one.” He imagines her to be just like that shimmering beauty he spotted that night in that silver dress at that concert two years ago. Zelda, having grown up with a hippie mom who believed the universe would provide for everything, rebelled into a no-nonsense practical lawyer who prefers the control of online dating. But when a computer glitch sends her a total mismatch, she’s asked to come in for an interview at the Internet dating site where Andrew works, and this is where it all begins. Andrew and Zelda meet for the first time and despite their differences, sparks fly. She thinks it’s chance. He thinks it’s fate. After all, he’s convinced she’s the shimmering girl in the silver dress. Is it true love forever or just a detour in destiny? When Andrew's ex-girlfriend Madeline (guest star Dreama Walker) crashes at Andrew's while she is town, Zelda fears she is secretly after Andrew. Feeling threatened, Zelda decides to set up Madeline with an ex-boyfriend of hers in hopes of distracting her. Adding to her stress, Zelda contemplates a major career move. When Andrew’s (Ben Feldman) father Pete (guest star Dan Lauria) unexpectedly shows up to visit, Zelda (Cristin Milioti) tries to win his affection after Pete tells Andrew he isn’t a fan of hers. Elsewhere, Big Bird (Christina Kirk) implements a new cancellation policy at Wallflower, despite not having Dane’s (guest star Wayne Wilderson) corporate approval. Between the constant distractions of Stu (Henry Zebrowski), Stephie (Lenora Crichlow) and her new boyfriend Joseph (guest star Echo Kellum), Andrew (Ben Feldman) and Zelda (Cristin Milioti) can’t seem to get in any alone time. This causes some friction and Andrew and Stu start to look for separate apartments. Meanwhile, Big Bird (Christina Kirk) recruits Lora (Hong Chau) for some girl talk as they strategize a way to get Dane (guest star Wayne Wilderson) to notice Big Bird. Zelda sets up Stu with a stenographer from her office, Jennifer (guest star SARAH BAKER - The Campaign), who Zelda feels would be perfect for Stu. To Andrew's surprise, everything is going seamlessly with the new couple - except Stephie feels like a fifth wheel. Andrew seeks help from Lora to match Stephie with someone at Wallflower so she doesn't feel left out. Andrew and Zelda try to pick out Christmas gifts for one another with help from their friends. Meanwhile, Big Bird faces a crisis when a customer signs up for a news interview about an unsuccessful matching experience.